As a freshman in 2009, Robert Lester played in eight games, recording 8 tackles. He played on special teams in the national championships game teams in the national championship game against Texas, but did not record a tackle. As a sophomore, Lester took over as a starter and had 52 tackles, 8 interceptions and a sack. As a junior, Lester started all 13 games at strong safety during the crimson tide's 2011 national championship season. He had 39 tackles, 3 pass breakups, a forced fumble, 1 tackle for loss, and a blocked field goal. He intercepted his first pass against Ole Miss and his second against LSU during the regular season.

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